diff --git a/.project/objectives/p3-27-biosphere-headless.md b/.project/objectives/p3-27-biosphere-headless.md
index 635c3500..9628c5e9 100644
--- a/.project/objectives/p3-27-biosphere-headless.md
+++ b/.project/objectives/p3-27-biosphere-headless.md
@@ -23,8 +23,7 @@ interact with.
 
 - [x] **Ecology population tick** ✅ (eb38e8678 + cba0ea8ec) — `process_ecology_phase` drives `EcologyEngine` per turn in `apply_end_turn` (relocated from mc-turn to dodge the mc-turn→mc-ecology→mc-mapgen cycle); seeds genesis on tick 1, persists via continuation-JSON on GameState; FFI `set_ecology_species_json` + harness `_apply_ecology_species`. mc-player-api 138/0.
 - [~] **Flora succession** — `EcologyEngine::process_step` already returns + applies `FloraTransition`s (subsumed in the ecology tick above). Confirm whether a separate `mc-flora::FloraEngine` pass is still needed or if process_step covers it.
-- [ ] **Marine ecology** — port `marine_harvest.gd` (372 lines: fish-stock depletion/
-  recovery, coral-reef bleaching, `ocean_dead_fraction` → climate) into Rust + tick it.
+- [x] **Marine ecology (core)** ✅ — per-tile fish-stock + coral-reef + mangrove→fish feedback already tick inside `EcologyEngine::process_step` (engine.rs:416/424), which runs in the headless ecology phase; `ocean_dead_fraction` updates via the climate phase (physics.rs:800). No live `MarineHarvestScript` remains — fully Rust. **Refinement:** the global catastrophic ocean-collapse `mc_ecology::ocean::tick_ocean_state` is unwired (only called in its own tests) — needs a populated BiomeTagRegistry (boot config) or an API switch to the built-in `has_tag` before it can tick headless.
 - [~] **Bio-targeting events** — fauna disease applier DONE (5890b1c15): EcologyEngine::apply_disease_events drives plague/pandemic/ecological mortality + canopy/tier loss from the boot-loaded events_config, struck in the ecology phase. Remaining: marine (fish/reef) + lair_kill_chance + o2_delta atmospheric pass.
 - [~] Deterministic from seed ✅ (cargo green, determinism test); headless e2e boot pending dylib rebuild + GUT boot proof.