diff --git a/src/game/engine/src/modules/ecology/ecosystem_simplified.gd b/src/game/engine/src/modules/ecology/ecosystem_simplified.gd
index 78759a76..d9240188 100644
--- a/src/game/engine/src/modules/ecology/ecosystem_simplified.gd
+++ b/src/game/engine/src/modules/ecology/ecosystem_simplified.gd
@@ -190,7 +190,11 @@ static func _score_to_tier(score: float) -> int:
 	return 1
 
 
-static func recompute_biomes(tiles: Array, w: int, h: int, last_canopy: PackedFloat32Array, last_temp: PackedFloat32Array, last_moisture: PackedFloat32Array) -> void:
+static func recompute_biomes(
+	tiles: Array, w: int, h: int,
+	last_canopy: PackedFloat32Array, last_temp: PackedFloat32Array,
+	last_moisture: PackedFloat32Array,
+) -> void:
 	## Reclassify biomes where canopy/temp/moisture changed significantly.
 	## Updates last_* arrays in-place for next turn's comparison.
 	var n: int = tiles.size()
@@ -214,7 +218,11 @@ static func recompute_biomes(tiles: Array, w: int, h: int, last_canopy: PackedFl
 		last_canopy[i] = tile.canopy_cover
 		last_temp[i] = tile.temperature
 		last_moisture[i] = tile.moisture
-		if d_canopy > BIOME_CANOPY_DELTA or d_temp > BIOME_TEMP_DELTA or d_moisture > BIOME_MOISTURE_DELTA:
+		if (
+			d_canopy > BIOME_CANOPY_DELTA
+			or d_temp > BIOME_TEMP_DELTA
+			or d_moisture > BIOME_MOISTURE_DELTA
+		):
 			# Inline classifier call — will be transpiled to classifyBiome(tile)
 			var new_biome: String = _classify_biome(tile)
 			if new_biome != tile.biome_id:
diff --git a/src/game/engine/src/modules/ecology/fauna.gd b/src/game/engine/src/modules/ecology/fauna.gd
index 2013ffa3..0e098c56 100644
--- a/src/game/engine/src/modules/ecology/fauna.gd
+++ b/src/game/engine/src/modules/ecology/fauna.gd
@@ -4,11 +4,12 @@ extends RefCounted
 ## fish reproduction, reef growth, habitat suitability, carrying capacity, migration.
 
 const HexUtilsScript = preload("res://engine/src/map/hex_utils.gd")
+const LandFaunaModelScript = preload("res://engine/src/models/world/land_fauna.gd")
 
+var rng: RandomNumberGenerator = RandomNumberGenerator.new()
 var _biome_regions: Dictionary = {}
 var _regions_dirty: bool = true
-var rng: RandomNumberGenerator = RandomNumberGenerator.new()
-var _lp: Variant = null  # cached LandFaunaModel for current turn
+var _lp: LandFaunaModelScript = null  # cached LandFaunaModel for current turn
 
 
 func invalidate_region_cache() -> void:
@@ -341,7 +342,12 @@ func _find_target(
 			elif not ct.is_water() and ct.biome_id != ob:
 				continue
 			if _is_whale(sp) and mp != null:
-				if not ct.is_water() or ct.depth_from_coast < mp.whale_min_depth or ct.depth_from_coast > mp.whale_max_depth or ct.quality < mp.whale_min_quality:
+				if (
+					not ct.is_water()
+					or ct.depth_from_coast < mp.whale_min_depth
+					or ct.depth_from_coast > mp.whale_max_depth
+					or ct.quality < mp.whale_min_quality
+				):
 					continue
 			var count: int = ecology_db.get_creature_count_on_tile(c.x, c.y)
 			var cap: float = sp.get("carrying_capacity", 10.0)
@@ -478,7 +484,11 @@ func _rebuild_biome_regions(game_map: Variant) -> void:
 # -- Helpers --
 
 func _is_whale(sp: Dictionary) -> bool:
-	return sp.get("size", "") == "huge" and sp.get("habitat", "") == "aquatic" and sp.get("diet", "") == "carnivore"
+	return (
+		sp.get("size", "") == "huge"
+		and sp.get("habitat", "") == "aquatic"
+		and sp.get("diet", "") == "carnivore"
+	)
 
 func _perturbation() -> float:
 	var p: float = _lp.perturbation if _lp != null else 0.05
diff --git a/src/game/engine/src/modules/ecology/flora.gd b/src/game/engine/src/modules/ecology/flora.gd
index 6232b57a..e15abeb5 100644
--- a/src/game/engine/src/modules/ecology/flora.gd
+++ b/src/game/engine/src/modules/ecology/flora.gd
@@ -28,7 +28,8 @@ var _des: Dictionary = {}
 var _params_loaded: bool = false
 var _regrowth_stages: Array = []
 
-# Pre-resolved biome data: biome_id -> {canopy, undergrowth, fungi, temp_min, temp_max, moist_min, moist_max}
+## Pre-resolved biome data: biome_id ->
+## {canopy, undergrowth, fungi, temp_min, temp_max, moist_min, moist_max}
 var _biome_flora: Dictionary = {}
 
 
@@ -81,7 +82,9 @@ func process_turn(game_map: RefCounted) -> void:
 # =========================================================================
 
 
-static func tick_pioneer(tiles: Array, biome_flora: Dictionary, veg: Dictionary, o2_fraction: float = 0.21) -> void:
+static func tick_pioneer(
+	tiles: Array, biome_flora: Dictionary, veg: Dictionary, o2_fraction: float = 0.21
+) -> void:
 	## Pioneer colonization: seeds bare ground with initial flora using raw climate values.
 	## Uses temperature/moisture directly (not biome climate range) because abiotic-classified
 	## tiles may have narrow climate ranges that don't match their actual climate conditions
@@ -120,7 +123,9 @@ static func tick_pioneer(tiles: Array, biome_flora: Dictionary, veg: Dictionary,
 			tile.canopy_cover = pioneer_rate * 0.5 * hab
 
 
-static func tick_canopy(tiles: Array, biome_flora: Dictionary, veg: Dictionary, o2_fraction: float = 0.21) -> void:
+static func tick_canopy(
+	tiles: Array, biome_flora: Dictionary, veg: Dictionary, o2_fraction: float = 0.21
+) -> void:
 	## Grow canopy toward biome climax. Decay when outside climate range.
 	## Requires existing population (dP/dt = r*P) and sufficient atmospheric O2.
 	var growth_rate: float = veg.get("growth_rate", 0.02)
@@ -149,7 +154,9 @@ static func tick_canopy(tiles: Array, biome_flora: Dictionary, veg: Dictionary,
 			tile.canopy_cover = maxf(tile.canopy_cover - decay_rate, 0.0)
 
 
-static func tick_undergrowth(tiles: Array, biome_flora: Dictionary, veg: Dictionary, o2_fraction: float = 0.21) -> void:
+static func tick_undergrowth(
+	tiles: Array, biome_flora: Dictionary, veg: Dictionary, o2_fraction: float = 0.21
+) -> void:
 	## Grow undergrowth, capped by canopy shade. Decays faster in drought.
 	## Requires existing population (dP/dt = r*P) and sufficient atmospheric O2.
 	var growth_rate: float = veg.get("growth_rate", 0.02)