diff --git a/engine/src/modules/ecology/ecosystem.gd b/engine/src/modules/ecology/ecosystem.gd
index 23663a4a..70dd90d1 100644
--- a/engine/src/modules/ecology/ecosystem.gd
+++ b/engine/src/modules/ecology/ecosystem.gd
@@ -314,4 +314,7 @@ static func _is_water(tile: Variant) -> bool:
 		return sub in [
 			"deep_water", "shallow_water", "lake_bed",
 		]
-	return tile.biome_id in ["ocean", "coast"]
+	return tile.biome_id in [
+		"ocean", "coast", "deep_ocean", "shallow_ocean", "coral_reef",
+		"estuary", "mangrove", "lake", "pond", "river", "inland_sea",
+	]
diff --git a/engine/src/modules/ecology/ecosystem_simplified.gd b/engine/src/modules/ecology/ecosystem_simplified.gd
index 4384aa3f..86cc7567 100644
--- a/engine/src/modules/ecology/ecosystem_simplified.gd
+++ b/engine/src/modules/ecology/ecosystem_simplified.gd
@@ -281,4 +281,7 @@ static func _is_water(tile: Variant) -> bool:
 	if "substrate_id" in tile:
 		var sub: String = tile.substrate_id
 		return sub in ["deep_water", "shallow_water", "lake_bed"]
-	return tile.biome_id in ["ocean", "coast"]
+	return tile.biome_id in [
+		"ocean", "coast", "deep_ocean", "shallow_ocean", "coral_reef",
+		"estuary", "mangrove", "lake", "pond", "river", "inland_sea",
+	]
diff --git a/engine/src/modules/ecology/fauna_simplified.gd b/engine/src/modules/ecology/fauna_simplified.gd
index e9843341..d701fdb3 100644
--- a/engine/src/modules/ecology/fauna_simplified.gd
+++ b/engine/src/modules/ecology/fauna_simplified.gd
@@ -116,7 +116,10 @@ static func _is_water(tile: Variant) -> bool:
 	if "substrate_id" in tile:
 		var sub: String = tile.substrate_id
 		return sub in ["deep_water", "shallow_water", "lake_bed"]
-	return tile.biome_id in ["ocean", "coast"]
+	return tile.biome_id in [
+		"ocean", "coast", "deep_ocean", "shallow_ocean", "coral_reef",
+		"estuary", "mangrove", "lake", "pond", "river", "inland_sea",
+	]
 
 
 static func _get_neighbor_offsets(col: int) -> Array:
diff --git a/engine/src/modules/ecology/flora.gd b/engine/src/modules/ecology/flora.gd
index 4e4e6b8b..9333d4e9 100644
--- a/engine/src/modules/ecology/flora.gd
+++ b/engine/src/modules/ecology/flora.gd
@@ -299,7 +299,10 @@ static func _is_water(tile: Variant) -> bool:
 	if "substrate_id" in tile:
 		var sub: String = tile.substrate_id
 		return sub in ["deep_water", "shallow_water", "lake_bed"]
-	return tile.biome_id in ["ocean", "coast"]
+	return tile.biome_id in [
+		"ocean", "coast", "deep_ocean", "shallow_ocean", "coral_reef",
+		"estuary", "mangrove", "lake", "pond", "river", "inland_sea",
+	]
 
 
 static func _climate_match_flat(tile: Variant, bf: Dictionary) -> float: